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Note: This is being posted here instead of the Episode III or Clone Wars forum since this information is not going to be revealed in the upcoming movie or the third season of the cartoon series, but rather is from a Star Wars expanded universe novel.
Grievous was originally an organic being of the alien race known as the Kaleesh, which was at war with an enemy species known as the Huk. The Republic supported the Huk, and so the Kaleesh ended up getting support from the Separatists and in particular the Intergalactic Banking Clan.
San Hill, leader of the Banking Clan, noticed that Grievous was both a brilliant strategist and an excellent fighter, and asked Grievous to lead the Separatist troops throughout the galaxy against the Republic. Grievous refused, saying that he only cared about the war between the Kaleesh and the Huk, not about the larger conflict between the Separatists and the Republic. Hill brought this to the attention of Count Dooku and Darth Sidious, but despite their offers Grievous still refused to lead the Separatist army.
So Sidious, Dooku, and Hill arranged to have a bomb planted on the shuttle Grievous was on, and blame it on the Republic. Grievous was mortally wounded in the crash, and his unconscious, dying body was taken to the planet Geonosis, where his brain, eyes, and other organs were implanted into a cyborg body forged by Geonosians that would compliment his natural reflexes.
It might have been his rage at the Republic, who he believed caused his shuttle crash; it might have been a feeling of gratitude toward the Separatists for saving his life and giving him a new and improved body; or it might have been the modification that Banking Clan scientists made to his brain while he was unconscious; but when Grievous woke up in his new body he was more than willing to take up Hill's offer and agree to lead the Separatist troops throughout the galaxy against the Republic.
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